Capped Langur, Trachypithecus pileatus
CAPPED LANGUR
Trachypithecus pileatus

Geographic Distribution and Habitat
The capped langur (Trachypithecus pileatus) is a monkey species found in Bangladesh, Bhutan, India, and Myanmar. Within these regions, they may inhabit and thrive in several different ecosystems, such as subtropical, broadleaf, deciduous, and bamboo forests.
Geographically, the range of capped langurs is characterized by some significant landmarks, including the Chindwin River in Myanmar, the Brahmaputra River in Bangladesh, and the Himalayas, a mountain range that spans through Pakistan, India, Nepal, Bhutan, and China. They inhabit areas with elevations between 32.8 to 9,842 feet (10 and 3,000 meters), but because their home range is limited to around 0.08 to 0.25 square miles (21 to 64 hectares), it is unlikely that these borders greatly affect troop movement.
In Southeast Asia, the average temperature ranges between 77 to 95 degrees Fahrenheit (25 to 35 degrees Celsius). High humidity levels are very common, which leads to warm weather all year round, despite having a rainy and dry season. The rainy season occurs between the months of May to October, with the average rainfall per month equaling about 7.87 inches (200 millimeters), while the dry season occurs between December and April.
According to recent studies, there are three recognized subspecies of capped langurs (Trachypithecus pileatus): the capped langur (Trachypithecus pileatus pileatus), the Buff-bellied capped langur (Trachypithecus pileatus brahma), and the Tenebrous capped langur (Trachypithecus pileatus tenebricus).
The capped langur (Trachypithecus pileatus pileatus) occurs in northeastern India and northwestern Myanmar. Their population is widely distributed across their country’s highland mountains at an elevation range of up to 9,842 feet (3,000 meters).
Buff-bellied capped langurs occur in northeastern India. Their home range covers land north of the Brahmaputra River, and on the east bank of the river Jia-Bhoreli.
Lastly, Tenebrous capped langurs occur in Bhutan and north-eastern India. They are found in the country’s Manas region, to the north of the Brahmaputra River, and between the river Jia-Bhoreli.

Size, Weight, and Lifespan
Capped langurs are a sexually dimorphic species, which means there are slight differences in appearance between males and females. Typically, males weigh between 22 to 31 pounds (10 to 14 kilograms), and females, who are smaller, weigh between 13 to 22 pounds (6 to 10 kilograms). Excluding their tails, both males and females can grow and reach a body length of about 27.56 inches (70 centimeters); their tails add an additional 19.7 to 27.56 inches (50 to 70 centimeters) to their overall size.
Like other langur species, in the wild, capped langurs live for up to 12 years.
Appearance
Named for the dark patch of hair on top of their heads that resembles a hat, capped langurs are an easily recognizable primate! Compared to other langur species, capped langurs are striking because their fur is a mixture of yellowish, orange, gray, brown, and black. Adults also have solid black faces and skin, which is accompanied by small, long noses and piercing orange eyes. Tufts of pale-orange hair protrude from their faces, which also cover their “c-shaped” ears.
In color, a capped langur’s tail is a combination of brown, gray, and black. Additionally, the further away the tail extends from the body, the darker the shades of fur become. Nearly the same length as its body, the tail roughly measures between 19.7 and 27.56 inches (50 to 70 centimeters). Despite this length, capped langurs are unable to use their tails for grasping, holding, or manipulating objects. Capped langurs do not have prehensile tails and instead use their tails for balance, which is especially important when they are climbing trees and navigating the forest.
At birth, infants are born with slight color variations compared to their parents. While adults have dark or black skin, infants have pale, cream-colored skin and orange fur. “Why is that?”, you may ask. Scientists and primatologists suggest that the genes responsible for orange fur in infants are ancestral, meaning they are the “original” gene from which modern genes have evolved. For many primates, including capped langurs, it is believed that an infant’s orange coloring helps adults to quickly locate their newborns quickly, which may also encourage greater care. Over the course of about 6 months, as they mature, an infant’s skin and fur will gradually darken until it closely resembles the coloring of an adult.

Diet
Capped langurs are mostly folivorous primates, meaning their diet mainly consists of leaves. According to a recent study, researchers discovered that 60% of a capped langur’s diet was made of leaves, and the average langur foraged for as many as 43 different plant species throughout its lifetime. In addition to leaves, they also consume other food products such as bark, seeds, flowers, fruits, or small insect species such as caterpillars. The favorite plant species of capped langurs are banyan fig trees (Ficus benghalensis), sacred figs (Ficus religiosa), hog plums (Spondias mangifera), bahedas (Terminalia bellerica), kamala trees (Mallotus philippensis), and hareins (Litsea polyantha).
Most of the water that a capped langur obtains is primarily from the moisture in their food, or directly from a tree source.
To help with their digestion, capped langurs have an evolved dental formula that promotes easier leaf and seed eating. Like other leaf-eating primates, capped langurs have a dental formula of 2:1:2:3, meaning they have two incisors, one canine, two premolars, and three molars in each quadrant of their mouth. Their teeth, particularly their molars, allow them to shear and break plant material, which is important because improperly torn plant material can be especially hard on their stomachs and intestines. Luckily, their bodies are well prepared!
In addition to having a unique dental formula, capped langurs have a specialized stomach that helps them to break down leaves for digestion. Plant material, like cellulose, can be very rough on an animal’s digestive system, so langurs are outfitted with digestive adaptations to properly break down these tough fibers. Their stomach, which is sacculated and multi-chambered, contains a variety of microflora, or bacteria, that ferment and break down ingested food and is particularly beneficial for consumed plant materials. How neat!
Behavior and Lifestyle
Capped langurs are a predominantly arboreal species and spend nearly their whole lives living in the forest. They are diurnal, so most of their daily activity occurs during the day. Sleep, or general periods of rest, occurs throughout the night, or after the sun has set.
In order to obtain energy and perform daily activities, langurs consume a significant amount of leaves, spend about 40% of their day feeding. In a process known as photosynthesis, plants can convert sunlight energy into chemical energy, or glucose. After ingestion, glucose, also known as sugar, is broken down by the body and used as a primary source of energy for various cell types, tissues, organs, etc. Unsurprisingly, this process can take a lot of time to complete. Furthermore, fiber, which is typically abundant in plant products, is considered very hard to digest. When fiber is not properly digested, it can lead to bloating, gas, abdominal pain, or constipation. To avoid these difficulties, langurs spend the majority of their day resting and caring for their digestive health, which is why their home range is relatively limited. Although these processes are time-consuming, it is better for a capped langur to be safe than sorry!
When capped langurs are not feeding or resting, they engage in various forms of social and non-social play. Examples of play include hopping, running, jumping, climbing, wrestling, and chasing. Although infants and juveniles often play the most, each technique may help an individual establish social bonds or help them to develop survival techniques, which for everyone, is essential.
Land predators of capped langurs include golden jackals (Canis aureus), clouded leopards (Neofelis nebulosa), and leopards (Panthera pardus). Avian predator species include large raptors like crested serpent eagles (Spilornis cheela) or mountain-hawk eagles (Nisaetus nipalensis).
In Hinduism, langurs are culturally cherished and believed to be a symbol of the monkey deity Hanuman, the most devoted follower of Prince Rama, a central figure who is considered the incarnation of the god Vishnu. According to religious text, Hanuman is regarded as a symbol of courage, strength, and selfless service and is depicted as a half-monkey, half-human.
Although Hanuman langurs (Semnopithecus entellus) are independently named after the Hindu god and believed to be direct descendants of Hanuman, others believe additional langur species share Hanuman’s ancestry. Many devout Hindus identify the black skin of Hanuman langurs as proof of their relationship to the monkey god, so because capped langurs share this feature, they would also be considered descendants.
As a sign of respect to Lord Hanuman, langurs are celebrated in Hinduism. In fact, many people, especially in urban environments, leave food and water around Hindu temples and shrines to show admiration and appreciation.
Capped langurs live in groups that consist of a dominant male, several females, and a handful of juveniles and infants. The average troop size is around 9 to 11 individuals, and the ratio of males to females in a troop is typically 1 male for every 7 females. Once mature, if two or more males live together in a troop, they will eventually compete for dominance through aggressive displays. Comparatively, females care for their troop’s infants and juveniles and instead develop close bonds with one another.
In the morning, troop members wake early but remain at their sleeping sites until the sun has fully risen. Once risen, they move to higher branches to bask in the early-morning sun and heat, before eventually venturing off to forage and feed. After splitting up into smaller sub-groups, when foraging, langurs may spread out over wide distances to maximize their feeding potential. To avoid getting lost, they use contact calls to communicate to one another and share locations.
Because the home ranges of capped langurs overlap with those of other primate species, it is not uncommon to find capped langurs interacting with other primates whilst foraging. Examples of such primates include Hoolock gibbons (Hoolock), Nepal gray langurs (Semnopithecus schistaceus), or Phayre’s langurs (Trachypithecus phayrei).
Although many of these interactions result in a sympatric relationship, in which different species may occupy the same geographic area without interbreeding, some interactions do not have positive outcomes. Male langurs are generally considered territorial and may produce aggressive displays when threatened, especially when they feel that their home range is in danger. For example, when two or more males encounter each other, they may leap towards intruders or rapidly jump from branch to branch to cause disruption and disturbance.
Unfortunately, there are other problems between langur species, particularly when it comes to species interbreeding. For example, in Bangladesh, a recent study revealed instances in which capped langurs and Phayre’s langurs bred and hybridized.
In biology, hybridization refers to the interbreeding of two genetically distinct individuals that results in an offspring containing traits from both parents. Interspecific hybridization refers to the interbreeding of two different species. For many wild species, hybridization is dangerous because it may lead to a loss of an individual species’ unique genetic traits. When small, isolated populations breed and hybridize, gene flow, or the movement of genes from one population to another, becomes limited. When gene flow is impeded in this way, individuals are likely to interbreed and form a new species through a process known as speciation. Increased speciation can lead to the extinction of one or more parent species, causing all involved to be vulnerable.
Unfortunately, there is no specific, universally recognized name for the offspring of Phayre’s and capped langurs. Although mixed species groups of capped langurs and Phayre’s langurs have been observed successfully interacting with their hybrid, the interaction will not be observable for years to come. Hybridized individuals are usually born infertile, which means they cannot produce viable offspring, so each generation of hybrids will probably only exist for around 12 years. Unfortunately, in Bangladesh specifically, national conservation laws only protect pure langurs, not hybrids, which puts the hybrid species at an even greater disadvantage.
Capped langurs have a variety of vocal calls and gestures used to communicate specific messages, such as whooping, barking, growling, coughing, peeping, and begging. Interestingly, studies have also identified that many of these calls are specific to different sex-age classes. For example, females, or juveniles, may emit a high-pitched call during conflict to communicate their distress, while adult males may produce a deeper, low-pitched growl to communicate a similar message.
In order to maintain social hierarchies and dominance within their troops, langurs groom each other in a process known as allogrooming. Allogrooming serves as a social-bonding behavior that maintains group cohesion. Like allogrooming, capped langurs also participate in auto grooming, which is an animal’s innate behavior to maintain its own body surface through activities such as chewing, scratching, licking, rubbing, etc. Both autogrooming and allogrooming have the psychological benefit of reducing an individual’s stress and anxiety levels, in addition to having hygienic benefits of keeping the fur clean and parasite-free.
Reproductively mature capped langurs breed during Southeast Asia’s dry season. After their 200-day gestation period (pregnancy), females give birth to one infant. This infant typically weighs around 1 pound (454 grams).
Once an infant is born, nearly all parental responsibilities are executed by females. Fortunately, within a troop, adult females participate in a social behavior known as allomothering. By definition, allomothering refers to the parental care provided by group members other than the genetic mothers. So, all troop females share the responsibilities of caring for each other’s infants, whether feeding, grooming, holding, nursing, or protecting. This occurs until an infant matures and becomes independent, or nearly self-sufficient.
Females reach sexual maturity around 3 to 4 years of age, while males reach maturity between the ages of 6 and 7 years of age.

Capped langurs are leaf-eating primates, so they play a large part in maintaining their ecosystem’s integrity.
In addition to plants, langurs consume various types of seeds and fruits and are therefore plant predators to different species across their home range. Each plant, fruit, or leaf that a langur consumes as sustenance has a seed. Whether the seed is tossed, dropped, or digested and excreted, capped langurs have the important responsibility of diversifying their ecosystem through seed germination. Seed germination, or the process by which a plant grows from a seed to a seedling, is crucial for a plant’s life cycle and population growth. Not only does seed germination ensure the continuation of a plant species, potentially across long distances, but it is the foundation for all plant growth.
Without animals like capped langurs, biodiverse ecosystems are at an increased risk, especially as civilizations around the world continue to undergo the processes of urbanization, mining, crop harvesting, industrial logging, etc.


The International Union for Conservation of Nature lists capped langurs as Vulnerable (IUCN, November 21, 2015), appearing on the IUCN Red List of Threatened Species.
Over 36 years (three generations), wild capped langur populations decreased by 30% because of habitat loss. Unfortunately, capped langur populations are expected to decline even further due to increased pressure for land development, unsustainable bamboo harvest, coal mining, limestone mining, meat trading, etc. Not only do these circumstances put individual animals at a greater risk, but they also increase the likelihood of habitat degradation and fragmentation, creating a significant loss in quality of habitat. Furthermore, poor law enforcement puts the conservation of capped langurs at an even greater risk.
Capped langurs are listed in Appendix I of the Convention on International Trade in Endangered Species (CITES), an international agreement between governments whose goal is to ensure that international trade in specimens of wild animals and plants does not threaten their survival.
According to the IUCN’s website, all South Asian populations of capped langurs are listed under Schedule I, Part I of the Indian Wildlife (Protection) Act, which was amended up to 2002. Populations in Bangladesh are protected under Schedule III of the Bangladesh Wildlife (Preservation) (Amendment) Act, 1974. In Myanmar, this species is protected under the Wildlife Protection Act.
Capped langurs are found in a large number of protected areas across their home range, including Nameri National Park and Pakhui Wildlife Sanctuary in India, Royal Manas National Park in Bhutan, etc.
- Yang L, Sun T, Zhou Y, Tang C, Huang C, Fan P, Zhou Q. Pattern of play behavior in infant (age 1 to 12 months) white-headed langurs in limestone forests, southwest China. Ecol Evol. 2022 Aug 4;12(8):e9160. doi: 10.1002/ece3.9160. Erratum in: Ecol Evol. 2022 Oct 11;12(10):e9420. doi: 10.1002/ece3.9420. PMID: 35949525; PMCID: PMC9351325.
- Craig, Timothy P. (2018). Ecological Speciation. obo in Evolutionary Biology. doi: 10.1093/obo/9780199941728-0116
- Das, J., Chetry, D., Choudhury, A.U. & Bleisch, W. 2024. Trachypithecus pileatus (amended version of 2020 assessment). The IUCN Red List of Threatened Species 2024: e.T22041A259357527. https://dx.doi.org/10.2305/IUCN.UK.2024-2.RLTS.T22041A259357527.en. Accessed on 26 March 2025.
- Anwaruddin Choudhury “Distribution and Current Status of the Capped Langur Trachypithecus pileatus in India, and a Review of Geographic Variation in its Subspecies,” Primate Conservation, 2014(28), 143-157, (1 December 2014)
- https://www.nationalgeographic.com/magazine/article/langur-monkeys#:~:text=Hindus%20revere%20them%20as%20a,the%20course%20of%20his%20heroism.
- https://www.eprc.asia/langurs/#:~:text=Despite%20having%20a%20long%20to,jumping%20amongst%20the%20forest%20canopy
- https://www.naturesafariindia.com/langur-monkey-species-in-india/#:~:text=Langurs%20are%20leaf%2Deating%20primates,to%20their%20distinctive%20large%20belly.
- https://www.nature.com/scitable/knowledge/library/primates-in-communities-the-ecology-of-competitive-59119961/
- https://greenverz.com/capped-langur/#:~:text=They%20also%20have%20white%20fur,habitat%20loss%20and%20other%20threats.
- https://www.inaturalist.org/taxa/43493-Trachypithecus-pileatus
- https://socialsci.libretexts.org/Courses/Cosumnes_River_College/ANTH_374%3A_Birth_to_Death_-The_Anthropology_of_Primate_Culture_and_Behavior_(Fortunato)/02%3A_Meet_the_primates_-_Evolution_Characteristics_and_Taxonomy/2.01%3A_Meet_the_Living_Primates/2.1.02%3A_Key_Traits_Used_to_Distinguish_Between_Primate_Taxa#:~:text=Leaves%20often%20have%20toxins%20in,tracts%20filled%20with%20specialized%20bacteria.
- https://economictimes.indiatimes.com/news/india/langurs-can-read-mind-use-eight-unique-begging-gestures-to-demand-food-from-people-study/articleshow/103571711.cms?from=mdr
- https://www.biologyonline.com/dictionary/hybridization#:~:text=It%20is%20sometimes%20used%20synonymously,(sometimes%2C%20even%20species).
- https://www.youtube.com/watch?v=Fc2KAOqKV_E
- https://news.mongabay.com/2024/10/langurs-in-bangladesh-face-extinction-as-hybridization-between-species-escalates/
- https://www.worldlandtrust.org/species/mammals/capped-langur/
- https://www.britannica.com/science/gestation
- https://www.wired.com/story/the-brutal-reason-why-some-primates-are-born-a-weird-color/#:~:text=The%20team’s%20hypothesis%20is%20that,more%20care%20from%20their%20mothers.
- https://blog.wcs.org/photo/2016/02/08/a-monkey-that-changes-color-lunar-new-year-bronx-zoo/#:~:text=It%20is%20believed%20that%20this%20outrageously%20orange,to%20the%20silvery%20color%20of%20the%20adults.
- https://brainmuseum.org/specimens/primates/langur/index.html#:~:text=Infants%20are%20weaned%20at%20about,Krishna%20Rivers%2C%20south%20of%20Ganges.
- https://www.nwf.org/Magazines/National-Wildlife/2003/Monkeys-From-Heaven#:~:text=Many%20devout%20Hindus%20regard%20the%20black%20skin,the%20living%20descendants%20of%20the%20monkey%20god.&text=%22The%20Hindus%20honor%20God%20Hanuman%20by%20provisioning,been%20studying%20Hanuman%20langurs%20for%20many%20years.
Written by Brooklynne Mitchell-Arno, March 2025