CENTRAL CHIMPANZEE

Pan troglodytes troglodytes

Geographic Distribution and Habitat

Central chimpanzees are found in the tropical and swamp forests of Central Africa, across Angola, Cameroon, Central African Republic, Congo, Democratic Republic of Congo (DRC), Equatorial Guinea, and Gabon (where they have been studied the most). The Sanaga River in Cameroon and the Congo River in the DRC act as natural barriers that prevent their movement to the north and south of their range.
These chimps typically live in lowland forests and prefer mixed tree species. However, due to extensive habitat destruction and change, they have adapted to foraging and nesting in forests dominated by single-tree species.

TAXONOMIC NOTES

The common chimpanzee (Pan troglodytes) is currently divided into four geographically separated subspecies: the Central (Pan troglodytes troglodytes), Eastern (Pan troglodytes schweinfurthii), Western (Pan troglodytes verus), and Nigerian-Cameroonian (Pan troglodytes ellioti) chimpanzees.

The taxonomical classification has been controversial, with some researchers arguing that the genetic differences between the subspecies are similar to the genetic variation found among humans. Therefore, the subspecies should be collapsed into a single species of chimpanzees. Multiple genetic analyses have shown that the differences between Central and Eastern chimpanzees are stable and have most likely accumulated over millions of years because of their geographical isolation. This indicates that subspecies classification is difficult but may be the most appropriate given our current knowledge.

Central chimpanzee geographic range. Map credit: Luís Fernández García/Creative Commons

Size, Weight, and Lifespan

Central chimpanzees are the largest of the chimpanzee subspecies, with males weighing an average of 132 pounds (59.7 kg) and females weighing an average of 101 pounds (45.8 kg). Males can reach a standing height of 5.2 feet (160 cm), while females can reach 4.3 feet (130 cm).

In the wild, their average lifespan is 15 to 25 years. In captivity, they can live to 40 to 50, on average. Their maximum lifespan can be 50-60 years. Chimpanzees can reach these old ages in the wild, but judging ages in the field isn’t easy. With the pressures of poaching and disease, these long lifespans seem optimistic in the wild

Appearance

All chimpanzees are tailless and have standard morphological features such as long dark fur, hairless faces with deep-set brown eyes, and their young are born with pale faces and a white tail tuft at the base of their spine, which darkens as they age. Central chimpanzees are larger and have significantly longer arms than other subspecies. They also have longer fingers and short thumbs, which indicate that Central chimps may be more suited to climbing and traveling through trees. Their faces also appear more prominent, with less hair than the other subspecies. 

Photo: © bureaubenjamin/iNaturalist/Creative Commons
Diet

Central chimpanzees, like all chimpanzee subspecies, are omnivores. Their diet consists of various foods, including vegetation, sap, honey, insects, and meat, but they primarily consume fruits. They also use tools to crack open palm nuts and collect insects. Although Central chimpanzees hunt and eat other animals, they engage in this behavior less frequently than other chimpanzee subspecies. In particular, they do not eat as many other primate species as the Eastern chimpanzee (Pan troglodytes schweinfurthii)

Their diets differ regionally and seasonally depending on the foods available. One region of Gabon has a population of Central chimps specializing in smashing tortoises against rocks and branches to eat the meat inside. Tortoises were hunted more in the dry season when they were more visible. These chimps even shared the meat with their party members.

Behavior and Lifestyle

Chimpanzees are diurnal, meaning they are active during the day. They spend most of their time in trees (arboreal), and their long arms and fingers give them the advantage of quickly swinging through the trees. They are comfortable walking terrestrially or on the ground. During the rainy season, they tend to spend more time in the canopy and travel more on ground during the dry season.  When terrestrial, they walk on all four limbs (quadrupedally), using their knuckles to support their weight. This ” knuckle-walking ” movement is the most efficient way to move on the ground because of their long arms, and they often can run in a bounding motion using this method. They can also walk upright on their legs (bipedally), but this is slower and more clumsy because their thigh bones do not angle out like humans’. 

Chimpanzees can live in large troops, but most of their activities and daily routines are done with smaller groups called parties. In most chimpanzee subspecies, the party males oversee territorial patrols and keep out intruders. Territories are defended by aggressive vocalizations, chasing, thrashing of vegetation, physical fighting, and even killings. Eastern chimps are famous for multi-year violent encounters and intergroup killings. Central chimpanzees are not recorded as having such long-term violent behaviors, even though there have been many instances of intergroup killings during short territorial disputes. Central chimpanzee intercommunity encounter rates are generally lower than other chimpanzee subspecies. Unlike Eastern chimpanzee troops, females and young Central chimpanzees sometimes participate in patrols and intergroup encounters. Having young offspring in the territorial patrols seems foolhardy because they are weaker and much more likely to get killed during encounters. This presence of the young on patrols may explain why encounter rates are low, as parties do not want to risk physical fights where the young are in danger of getting killed. When encounters occur in these communities, they are usually acoustic (yelling at each other) or visual aggressions without violence. 

Tool use has been well documented in Central chimpanzees. They often use sticks to dip into termite or ant nests, pull out the little insects, and eat them. This behavior is called dipping or fishing. Central chimpanzees sometimes carry multiple tools for between 33 feet and 230 feet (10 and 70 m) in preparation for ant-dipping opportunities. This amazing behavior suggests that chimpanzees have some planning capabilities, an initial part of complex problem-solving seen in humans. Mature chimpanzees transport tools more often than younger ones, suggesting they learn through experience. Many primates and birds have been documented as being able to manipulate or change another environmental condition using an external object. However, anticipating the need for tools, modifying existing tools, and teaching tools to others, like chimps, is considered a complex mental process unique to humans. 

Some Central chimpanzees select variations of an object in their environment that best suit their purpose and then modify these tools to make them more effective. For example, to eat termites from an above-ground termite nest, chimps use thinner herbaceous twigs to probe the natural exit holes. For underground termite nests, chimps use thicker branches to make a tunnel that reaches the nest before using sticks as probes. In either case, they chew on the end of the probe stick to make the ends more like a brush that can collect more insects in each scoop. The thicker woody sticks are more durable than the thin twigs, so they are often reused multiple times.

They also gather honey from beehives using sticks, often using longer sticks for bees with stingers and sturdy branches to break open hives of underground bees that do not have stingers. 

Tool use in chimps varies by community and region. For example, tortoise smashing is common in areas with more tortoises, while digging up underground bee nests is seen in other chip communities where they have learned to look for signs of these nests and dig them up to get grubs and honey. 

Chimpanzees learn through trial and error (experiential learning) and also from watching other chimps (social learning). Thus, a behavior such as ant-dipping or tortoise smashing is learned and passed on to the next generation of chimps within a community. Transferring these behaviors across regions occurs when chimps come together in their fission-fusion society or when females leave their natal group and join another party. Many scientists think this transference of behaviors along generations of chimpanzees indicates chimpanzee culture, similar to rituals in human cultures. 

All chimp species build smaller, rough nests for short daytime rests and more elaborate, comfortable nests for sleeping at night. Nighttime nests are made from bent branches shaped to a form base and then topped with soft ferns and leaves. These elaborate nests are commonly built in trees because they offer the most protection against predators. In general, ground nests are quickly constructed to rest during the day in between feeding bouts. Nighttime ground nesting or ground night nests are less common in Central chimpanzees than in other subspecies; only 4% of Central chimpanzee nests have been recorded as ground nests. They prefer ground night nesting in swamps during the dry season (when water levels are low and nesting sites are more available) because human hunters tend to avoid difficult-to-navigate swampy regions.  Researchers think that nesting may be an essential precursor to the evolution of human memory and cognition (ability to think). Sleeping in a safe and comfortable nest improves the length and quality of sleep. Deep sleep increases the release of hormones that establish memories and neuron (nerve cell) health. Therefore, nesting behavior that would have evolved among primates may have contributed to complex neural pathways found in modern primate brains. 

Chimpanzees were used in research for decades, and many of their behaviors have been studied under laboratory conditions. These laboratory studies help understand individual behaviors. However, they are challenging to apply to wild conditions where the environment, food availability, and presence of other animals significantly influence chimpanzee behavior. One interesting outcome of lab behavior research is understanding how chimpanzees use their gestures and eye contact to communicate across species, to humans specifically. Many studies have shown that chimpanzees use pointing and alternating gazing between objects and humans to communicate the object’s location to their human handlers. This behavior has also been noted in human babies, highlighting the common bases of language and contextual communications in humans and apes.

Fun Facts

Central chimpanzees are the largest of the common chimpanzee subspecies. 

They have longer fingers and shorter thumbs, making them best suited for arboreal life compared to other subspecies.

Females and young Central chimps sometimes join males during territorial patrols. 

They are sympatric (cohabit) with another great ape, the western lowland gorilla. 

Daily Life and Group Dynamics

Central chimpanzees form fission-fusion communities of over 60 individuals, consisting of multiple adult females, males, and their offspring. These large groups form when there are abundant food resources and fewer predators. They break into smaller groups called parties to travel and forage for food. Chimpanzees primarily spend their time in party groups. Party size can vary between 2 and 30 individuals, and members often leave and join depending on foraging needs and social interactions.

In chimpanzee societies, males and females have a separate dominance hierarchy, and males always outrank females. Male dominance is usually established linearly, with the strongest chimp taking the lead or “alpha’ position. However, other males will continuously fight for this position because it gives the ‘alpha’ male access to reproductive females and the best chance of passing his genes to the next generation. Contests among males are usually aggressive and noisy, and they even result in bloodshed. Female dominance contests are over access to the best food resources, where the older females get the first choice of the ripest fruits. Female contests are usually peaceful, and food sources are generally shared. In Central chimpanzees, ‘alpha’ females can play a significant role in chasing out intruding females because they travel with males on territorial patrols. 

Chimpanzees wake at dawn, and as a group, they leave their nests soon after to forage for food. Most of their day (about 50% of their active time) is spent feeding. They feed for about 2-3 hours in the morning and again before retiring for the night. Between their feeding bouts, around the hot midday period, they spend about 18-25% of their active time. They build simple resting nests for this rest period. These nests are usually in trees during the wet season and on the ground during the dry season. During these rest times, they also participate in social activities like playing and grooming each other. 14% of their time is spent moving, searching for food, or patrolling territorial boundaries. During the dry season, when fruits are less abundant, more time is spent searching for food and eating, and less time is spent resting. Central chimpanzees can travel long distances through dense forests, about 1-3 miles (2 to 5 km) daily. They usually find their sleeping sites and start constructing comfortable night nests about an hour before sunset. 

Most of these activities are done within the chimpanzee’s home range, between 10 and 22 square miles (27 to 59 square km). A smaller core territory is actively defended from neighboring communities, which helps the party maintain access to important fruiting and nesting trees. 

Communication

Chimpanzees use vocalizations to communicate social and ecological conditions to each other. For example, they make alarm calls when they see a predator and grunt to tell others that food is nearby. Scientists have identified 13 call types (e.g., pants, grunts, hoos)and many subtypes (e.g., pany hoots, alarm-hoos, rest-hoos) in chimpanzees. 

Hoos are quiet sounds made when chimpanzees make their night nests and settle in to sleep. 

Pant grunts and pant-barks are associated with social contexts such as greeting each other or a female ready to mate. The volume and length of these calls depend on the social dynamics between individuals. So if a dominant male is particularly aggressive, pant grunts from subordinate chimps are usually louder. 

Screams attract attention, but chimpanzees also modulate their screams to trigger different responses in each other. When an aggressor attacks, the victim uses a high-severity, loud-volume scream that triggers more attention from other chimps. The listener of the scream may decide to intervene depending on the social context of the situation; for example, a mother may intervene if her child is the victim.  

Most vocalizations are used in the early morning while they feed and in the evening when they gather to sleep. Chimp parties can be spread across the forest as they forage for food. Vocalizations help chimpanzees maintain contact without seeing each other, and they can also tell each other where a food source is. Chimpanzees emit a rough grunt when they find a suitable fruiting tree for the party, and the pitch and structure of the rough grunt change with the size of the tree. So, when a chimpanzee’s vocalizations indicate the presence of larger trees with more fruits, more of their party will be triggered to join the vocalizing chimp.  

In addition to vocalizations, Central chimpanzees have been documented using eye-gaze as an intentional communication method, especially for difficult-to-detect threats like snakes. When chimpanzees hear or see a snake, they make alarm calls and alternatively glance between other chimps and the snake’s location. In this case, vocal calls and physical gesturing combined are better at communicating the presence of danger to others. 

Gestures and hand signals, such as pointing and waving their arms, generally draw attention to communicate danger or aggression. In Central chimpanzees, these physical movements and aggressive vocalizations can ward off intruders and prevent the escalation of territorial encounters into violent events. 

Drumming is a non-vocal acoustic behavior in which chimpanzees beat tree trunks to make loud rhythmic sounds. It is used for long-distance communication, mostly in small groups that are more spread out. The drumming rhythm changes for each drummer, leading many to think that individuals have drumming signatures that communicate the identity of the drummer to the listener. Drumming is often done by males and done in conjunction with pant-hoots. 

Olfactory signals through scent secretions are a non-visual method of communication. Most of these signals come through urine and feces with hormones or gland secretions and contain information about the depositor. For example, females in estrus secrete reproductive hormones like estrogen in their urine, which attracts the attention of males and lets them know that they are ready to mate.

Reproduction and Family

Chimps mate and reproduce throughout the year. Both males and females reach puberty at around 7 to 8 years old. When a female is hormonally ready to mate (a condition called estrus), the skin around her reproductive organs (the anogenital region) swells. This conspicuous visible sign triggers males to initiate mating with the female. 

Mating patterns are variable. Females can be promiscuous and copulate (mate) with multiple males where there is no aggressive competition between males. She can also form consortships where one male gets exclusive mating access, and they leave the group during the mating period. There is also a possessive mating pattern where one male aggressively fights off other males to mate with the female. Mating pattern choice could be dependent on the number of reproductive females in the party (fewer females could trigger more possessive mating systems) or the number of males (fewer males means less mating competition and more tolerance towards peaceful promiscuous copulations).

The gestation period (length of pregnancy) is about 7.5 months. Usually, a single infant is born to the mother, though twins have been reported on rare occasions. The infant death rate is high. Only one-third of all the chimpanzees a female gives birth to will survive past infancy.

Chimpanzees take a long time to develop, and infants can depend on their mothers for nutrition for up to 5 years. Mothers are too busy nursing and taking care of their young, and usually do not get pregnant for about 6 years after they give birth. Adult females remain reproductive well into their forties. With high infant mortality and long inter-birth intervals, a female Central chimpanzee can have about nine offspring in her lifetime. The slow development, low infant survival, and low birth rates of chimpanzees make it difficult for a population to recover once their numbers have been reduced. 

A young chimp is considered a juvenile when they reach 5 to 7 years old. At this age, they play, forage away from their mothers, and interact with other party members. They are considered adolescents between the ages of 7 and 12. Females tend to mature faster and are considered adults by the time they become 13 years old. In comparison, male adults are usually about 15 years old. Males stay with their natal party (where they were born). Females disperse or leave their natal group to join a party in another region. This behavior prevents inbreeding between females and closely related chimps (uncles or cousins). Female dispersal is also a way of transferring behaviors across space. She takes what she has learned about tool use and foraging from her natal troop and teaches other chimps in her new family. 

Olga Chimp. Photo courtesy of © HELP-Congo. Used with permission
Ecological Role

Central chimpanzees are excellent seed dispersers. As frugivores, they travel large distances through the forest. They move seeds away from their parent plant by discarding them after eating the fruit or depositing them in their feces. Thus, they contribute to regenerating valuable fruiting trees in the forest ecosystem.

Central chimpanzees share their habitat with another great ape, the western lowland gorilla (Gorilla gorilla gorilla), or in other words, they are sympatric species. They also share many of the same food sources; their diets overlap 50 to 80% in some regions. In such cases, how do these great apes avoid competition and survive without pushing one species out of the habitat? Chimpanzees eat fruit throughout the year, and their digestive systems are better adapted to eating fat-rich fruits and nuts (such as coconuts). Gorillas have broader and stronger teeth, allowing them to eat more woody and leafy foods when fruits are unavailable. This slight separation in food choices or niche partitioning allows gorillas and chimpanzees to co-exist without competing.

As charismatic flagship species in Central Africa, Central chimpanzees have an intrinsic value where they function in maintaining a healthy ecosystem. Chimps need various fruiting trees to have enough food and avoid competition with other animals. Trees need chimps to disperse their seeds in areas with less competition from similar plants. Other animals benefit from the regeneration of these trees. Leopards prey on chimpanzees, while chimpanzees hunt smaller prey when fruits are not in season. Chimpanzees live in a complex system of animal and plant communities that interact to balance the ecosystem’s energetic needs.

Conservation Status and Threats

The International Union for Conservation of Nature lists the Central chimpanzee as Endangered (IUCN, 2016), appearing on the IUCN Red List of Threatened Species.

Central Africa is a hotspot for poaching, and illegal killing is the main reason for chimpanzee population declines in this region. Central chimpanzees are often found in bushmeat markets. The increase of logging and mining industries in previously untouched forests has increased road networks and access to wildlife, making hunting easier.

Human activities are invariably associated with deforestation, which negatively affects chimpanzee survival. Habitat changes remove important fruit trees. Palm oil plantations especially convert multi-species forests into monocultures that cannot provide food or adequate shelter for most wildlife, including chimps. The movement of more humans, agriculture, and development into chimpanzee habitats brings associated pollution of water and soils, which affects all wildlife.

Human movement and the introduction of foreign elements into a once-pristine habitat have increased diseases among chimpanzees. The most famous case of this was the deadly Ebola virus outbreak of the 1990s, which caused the death of hundreds of chimpanzees and gorillas in the Gabon-Congo area. Fruit bats are thought to be the source of the Ebola virus that was introduced to human systems when the bats were hunted for food. The proximity between humans and chimpanzees exposed the infected blood and secretions to wildlife and quickly transmitted through the primate populations. The Ebola virus spread between gorillas, chimpanzees, and humans. The simian immunodeficiency virus is the precursor to HIV (Human Immunodeficiency Virus) and is another zoonotic disease (that can be transmitted from animals to humans) that jumped to humans through contact with infected blood when Central chimpanzees were hunted and butchered.

Even a slight reduction in the natural population of chimpanzees can have long-term harmful effects on the species. These highly social primates depend on their large communities to find food and survive. They must also survive a high infant mortality rate and a long development period before a young chimp becomes an adult and ready to mate. Removal of chimpanzees from the system means fewer individuals will be available to mate and replenish the next generation of chimpanzees, and the generational gap will most likely become longer as the remaining chimpanzees have to survive in smaller groups.

Conservation Efforts

The Central chimpanzee is listed in Appendix I of the Convention on International Trade in Endangered Species (CITES), an international agreement between governments whose goal is to ensure that international trade in specimens of wild animals and plants does not threaten their survival.

All chimpanzees are legally protected from hunting and poaching, and trading any part of a chimpanzee is illegal. Despite this, enforcement and protection of chimpanzee populations are insufficient across their range. Protection agencies are usually understaffed and underfunded. Only 33% of Central chimpanzees are estimated to be protected, and only 22% of their range is actively patrolled by forest rangers. 

Initiatives to protect Central chimpanzees include increasing law enforcement agents in currently unprotected areas, preventing the clearing and further logging of large tracts of forest lands, and establishing international protection agreements in habitats that cross political boundaries. Disease transmission awareness and education efforts among local communities have also been implemented to prevent zoonotic outbreaks. 

Protecting large areas of forests from development offers the best chance of protecting existing chimpanzee populations. National parks protect areas in individual countries, but these are fragmented habitats. Many of the actions proposed by primate conservation groups aim to coordinate international coordination to connect protected habitats and coordinate protections against poaching. 

Humans value chimpanzees for their charismatic, human-like behaviors, which have created tourism and monetary value for their existence in the wild. As chimpanzee populations decline, it is ideal for governments, conservation organizations, and tourism industries to align sustainable tourism activities with preserving the wild behaviors and safety of chimpanzees in their natural environment. The IUCN Action Plan for Chimpanzees includes tourism activities that do not exploit chimpanzee habitats and populations and prioritize minimal dangerous contact (which can potentially transmit diseases) between chimps and humans. 

References:
  • Cawthon Lang KA. 2006 April 13. Primate Factsheets: Chimpanzee (Pan troglodytes) Taxonomy, Morphology, & Ecology . <http://pin.primate.wisc.edu/factsheets/entry/chimpanzee>.
  • Collins, E., Weladji, R.B. Occupancy of Sympatric Central Chimpanzee (Pan troglodytes troglodytes) and Western Lowland Gorilla (Gorilla gorilla gorilla) in Campo Ma’an Conservation Area, Southern Cameroon. Int J Primatol 45, 1264–1285 (2024). https://doi.org/10.1007/s10764-024-00446-2
  • Drakulovski, P., Bertout, S., Locatelli, S., Butel, C., Pion, S., Mpoudi-Ngole, E., Delaporte, E., Peeters, M. and Mallié, M., 2014. Assessment of gastrointestinal parasites in wild chimpanzees (Pan troglodytes troglodytes) in southeast Cameroon. Parasitology research, 113(7), pp.2541-2550.
  • Eleuteri, V., Henderson, M., Soldati, A., Badihi, G., Zuberbühler, K., & Hobaiter, C. (2022). The form and function of chimpanzee buttress drumming. Animal Behaviour, 192, 189-205. https://doi.org/10.1016/j.anbehav.2022.07.013
  • Ella, G.W.E., Takenoshita, Y., Koumba, L.B.M., Nguelet, F.L.M., Iwata, Y., Ando, C., Mavoungou, J.F. and Abernethy, K., 2023. Diet composition and feeding ecology of chimpanzees (Pan troglodytes troglodytes) in a lowland tropical forest in Moukalaba-Doudou National Park, Gabon. African Study Monographs, 43, pp.42-60.
  • Estienne, V., Stephens, C. and Boesch, C., 2017. Extraction of honey from underground bee nests by central African chimpanzees (Pan troglodytes troglodytes) in Loango National Park, Gabon: techniques and individual differences. American Journal of Primatology, 79(8), p.e22672.
  • Farmer, K.H., Buchanan-Smith, H.M. and Jamart, A., 2006. Behavioral adaptation of Pan troglodytes troglodytes. International Journal of Primatology, 27, pp.747-765.
  • Fünfstück, T., Arandjelovic, M., Morgan, D.B., Sanz, C., Reed, P., Olson, S.H., Cameron, K., Ondzie, A., Peeters, M. and Vigilant, L., 2015. The sampling scheme matters: Pan troglodytes troglodytes and P. t. schweinfurthii are characterized by clinal genetic variation rather than a strong subspecies break. American Journal of Physical Anthropology, 156(2), pp.181-191.
  • Funkhouser, J. A., Musgrave, S., Morgan, D., Kialiema, S. N., Ngoteni, D., Brogan, S., McElmurray, P., & Sanz, C. (2024). Chimpanzees employ context-specific behavioral strategies within fission–fusion societies. Primates; Journal of Primatology, 65(6), 541. https://doi.org/10.1007/s10329-024-01165-1
  • Head, J.S., Boesch, C., Makaga, L. and Robbins, M.M., 2011. Sympatric chimpanzees (Pan troglodytes troglodytes) and gorillas (Gorilla gorilla gorilla) in Loango National Park, Gabon: dietary composition, seasonality, and intersite comparisons. International Journal of Primatology, 32, pp.755-775.
  • Humle, T., Maisels, F., Oates, J.F., Plumptre, A. & Williamson, E.A. 2016. Pan troglodytes (errata version published in 2018). The IUCN Red List of Threatened Species 2016: e.T15933A129038584. https://dx.doi.org/10.2305/IUCN.UK.2016-2.RLTS.T15933A17964454.en. Accessed on 23 March 2025.
  • iNaturalist. Available from https://www.inaturalist.org/taxa/147737-Pan-troglodytes-troglodytes Accessed 3/23/2025
  • Jones, C., Jones, C.A., Jones Jr, J.K., Wilson, D.E. and Dken, P., MAMMALIAN SPECIES N0. 529, pp. 1-9, 3 figs.
  • Jones, C., 1971. Comparative ecology of Gorilla gorilla (Savage and Wyman) and Pan troglodytes (Blumembach) in Rio Muni, West Africa. Bibl Primatol, 13, pp.1-95.
  • Krause, M. A., & Fouts, R. S. (1997). Chimpanzee (Pan troglodytes) pointing: Hand shapes, accuracy, and the role of eye gaze. Journal of Comparative Psychology, 111(4), 330–336. https://doi.org/10.1037/0735-7036.111.4.330
  • Maisels, F., Strindberg, S., Greer, D., Jeffery, K., Morgan, D.L. & Sanz, C. 2016. Pan troglodytes ssp. troglodytes. The IUCN Red List of Threatened Species 2016: e.T15936A102332276.
  • Martínez-Íñigo, L., Baas, P., Klein, H., Pika, S. and Deschner, T., 2021. Intercommunity interactions and killings in central chimpanzees (Pan troglodytes troglodytes) from Loango National Park, Gabon. Primates, 62(5), pp.709-722.
  • Martínez-Íñigo, L., Baas, P., Klein, H., Pika, S., & Deschner, T. (2021). Home range size in central chimpanzees (Pan troglodytes troglodytes) from Loango National Park, Gabon. Primates; Journal of Primatology, 62(5), 723. https://doi.org/10.1007/s10329-021-00927-5
  • Mittermeier, R.A., Rylands, A.B. and Wilson D.E. 2013. Handbook of the Mammals of the World: Volume 3 Primates. Lynx Edicions, Barcelona.
  • Musgrave, S., Koni, D., Morgan, D. and Sanz, C., 2024. Planning abilities of wild chimpanzees (Pan troglodytes troglodytes) in tool-using contexts. Primates, 65(6), pp.525-539.
  • Pika, S., Klein, H., Bunel, S., Baas, P., Théleste, E. and Deschner, T., 2019. Wild chimpanzees (Pan troglodytes troglodytes) exploit tortoises (Kinixys erosa) via percussive technology. Scientific reports, 9(1), p.7661.
  • Slocombe, K. E., Lahiff, N. J., Wilke, C., & Townsend, S. W. (2022). Chimpanzee vocal communication: What we know from the wild. Current Opinion in Behavioral Sciences, 46, 101171. https://doi.org/10.1016/j.cobeha.2022.101171
  • Tagg, N., Willie, J., Petre, C. A., & Haggis, O. (2013). Ground Night Nesting in Chimpanzees: New Insights from Central Chimpanzees (Pan troglodytes troglodytes) in South-East Cameroon. Folia Primatologica, 84(6), 362-383.
  • UNGRASP. https://www.un-grasp.org/great-apes/chimpanzees. accessed March 26, 2025.

Written by Acima Cherian, March 2025